diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/RootMotion/SkeletonRootMotionBase.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/RootMotion/SkeletonRootMotionBase.cs
index c4fb7039b..a85b077ba 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/RootMotion/SkeletonRootMotionBase.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/RootMotion/SkeletonRootMotionBase.cs
@@ -213,7 +213,7 @@ namespace Spine.Unity {
 				Vector2 parentBoneScale;
 				GetScaleAffectingRootMotion(out parentBoneScale);
 				ClearEffectiveBoneOffsets(parentBoneScale);
-				skeletonComponent.Skeleton.UpdateWorldTransform(Skeleton.Physics.Update);
+				skeletonComponent.Skeleton.UpdateWorldTransform(Skeleton.Physics.Pose);
 			}
 			ClearRigidbodyTempMovement();
 
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
index 032dd46f4..ed1c6a4dd 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
@@ -83,10 +83,15 @@ namespace Spine.Unity {
 		/// </summary>
 		public event UpdateBonesDelegate UpdateLocal { add { _UpdateLocal += value; } remove { _UpdateLocal -= value; } }
 
-		/// <summary>
+		/// <summary><para>
 		/// Occurs after the Skeleton's bone world space values are resolved (including all constraints).
 		/// Using this callback will cause the world space values to be solved an extra time.
-		/// Use this callback if want to use bone world space values, and also set bone local values.</summary>
+		/// Use this callback if want to use bone world space values, and also set bone local values.
+		/// </para><para>
+		/// When used in combination with PhysicsConstraints at your skeleton, you might want to consider adjusting
+		/// <see cref="MainPhysicsUpdate"/> and <see cref="AdditionalPhysicsUpdate"/> to save updates by setting one to
+		/// <see cref="Skeleton.Physics.Pose"/>.
+		/// </para></summary>
 		public event UpdateBonesDelegate UpdateWorld { add { _UpdateWorld += value; } remove { _UpdateWorld -= value; } }
 
 		/// <summary>
@@ -104,6 +109,15 @@ namespace Spine.Unity {
 		/// Instance SkeletonAnimation.timeScale will still be applied.</summary>
 		[SerializeField] protected bool unscaledTime;
 		public bool UnscaledTime { get { return unscaledTime; } set { unscaledTime = value; } }
+
+		protected Skeleton.Physics mainPhysicsUpdate = Skeleton.Physics.Update;
+		protected Skeleton.Physics additionalPhysicsUpdate = Skeleton.Physics.Update;
+		/// <summary>Physics update mode used in the main call to skeleton.UpdateWorldTransform().</summary>
+		public Skeleton.Physics MainPhysicsUpdate { get { return mainPhysicsUpdate; } set { mainPhysicsUpdate = value; } }
+		/// <summary>Physics update mode used at optional additional calls to skeleton.UpdateWorldTransform(),
+		/// such as after the <see cref="UpdateWorld"/> callback if a method is subscribed at the UpdateWorld delegate.
+		/// </summary>
+		public Skeleton.Physics AdditionalPhysicsUpdate { get { return additionalPhysicsUpdate; } set { additionalPhysicsUpdate = value; } }
 		#endregion
 
 		#region Serialized state and Beginner API
@@ -264,11 +278,11 @@ namespace Spine.Unity {
 			if (_UpdateLocal != null)
 				_UpdateLocal(this);
 
-			UpdateWorldTransform();
+			UpdateWorldTransform(mainPhysicsUpdate);
 
 			if (_UpdateWorld != null) {
 				_UpdateWorld(this);
-				UpdateWorldTransform();
+				UpdateWorldTransform(additionalPhysicsUpdate);
 			}
 
 			if (_UpdateComplete != null) {
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
index 643fbf179..828e13ed5 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
@@ -403,19 +403,19 @@ namespace Spine.Unity {
 			if (UpdateLocal != null)
 				UpdateLocal(this);
 
-			UpdateWorldTransform();
+			UpdateWorldTransform(mainPhysicsUpdate);
 
 			if (UpdateWorld != null) {
 				UpdateWorld(this);
-				UpdateWorldTransform();
+				UpdateWorldTransform(additionalPhysicsUpdate);
 			}
 
 			if (UpdateComplete != null)
 				UpdateComplete(this);
 		}
 
-		protected void UpdateWorldTransform () {
-			skeleton.UpdateWorldTransform(Skeleton.Physics.Update);
+		protected void UpdateWorldTransform (Skeleton.Physics physics) {
+			skeleton.UpdateWorldTransform(physics);
 		}
 
 		public void LateUpdate () {
@@ -644,6 +644,15 @@ namespace Spine.Unity {
 		[SerializeField] protected bool unscaledTime;
 		public bool UnscaledTime { get { return unscaledTime; } set { unscaledTime = value; } }
 
+		protected Skeleton.Physics mainPhysicsUpdate = Skeleton.Physics.Update;
+		protected Skeleton.Physics additionalPhysicsUpdate = Skeleton.Physics.Update;
+		/// <summary>Physics update mode used in the main call to skeleton.UpdateWorldTransform().</summary>
+		public Skeleton.Physics MainPhysicsUpdate { get { return mainPhysicsUpdate; } set { mainPhysicsUpdate = value; } }
+		/// <summary>Physics update mode used at optional additional calls to skeleton.UpdateWorldTransform(),
+		/// such as after the <see cref="UpdateWorld"/> callback if a method is subscribed at the UpdateWorld delegate.
+		/// </summary>
+		public Skeleton.Physics AdditionalPhysicsUpdate { get { return additionalPhysicsUpdate; } set { additionalPhysicsUpdate = value; } }
+
 		/// <summary> Occurs after the vertex data populated every frame, before the vertices are pushed into the mesh.</summary>
 		public event Spine.Unity.MeshGeneratorDelegate OnPostProcessVertices;
 
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
index ef193c2a9..5a2d35f88 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
@@ -63,10 +63,15 @@ namespace Spine.Unity {
 		/// Use this callback when you want to set bone local values.</summary>
 		public event UpdateBonesDelegate UpdateLocal { add { _UpdateLocal += value; } remove { _UpdateLocal -= value; } }
 
-		/// <summary>
+		/// <summary><para>
 		/// Occurs after the Skeleton's bone world space values are resolved (including all constraints).
 		/// Using this callback will cause the world space values to be solved an extra time.
-		/// Use this callback if want to use bone world space values, and also set bone local values.</summary>
+		/// Use this callback if want to use bone world space values, and also set bone local values.
+		/// </para><para>
+		/// When used in combination with PhysicsConstraints at your skeleton, you might want to consider adjusting
+		/// <see cref="MainPhysicsUpdate"/> and <see cref="AdditionalPhysicsUpdate"/> to save updates by setting one to
+		/// <see cref="Skeleton.Physics.Pose"/>.
+		/// </para></summary>
 		public event UpdateBonesDelegate UpdateWorld { add { _UpdateWorld += value; } remove { _UpdateWorld -= value; } }
 
 		/// <summary>
@@ -77,6 +82,15 @@ namespace Spine.Unity {
 
 		[SerializeField] protected UpdateTiming updateTiming = UpdateTiming.InUpdate;
 		public UpdateTiming UpdateTiming { get { return updateTiming; } set { updateTiming = value; } }
+
+		protected Skeleton.Physics mainPhysicsUpdate = Skeleton.Physics.Update;
+		protected Skeleton.Physics additionalPhysicsUpdate = Skeleton.Physics.Update;
+		/// <summary>Physics update mode used in the main call to skeleton.UpdateWorldTransform().</summary>
+		public Skeleton.Physics MainPhysicsUpdate { get { return mainPhysicsUpdate; } set { mainPhysicsUpdate = value; } }
+		/// <summary>Physics update mode used at optional additional calls to skeleton.UpdateWorldTransform(),
+		/// such as after the <see cref="UpdateWorld"/> callback if a method is subscribed at the UpdateWorld delegate.
+		/// </summary>
+		public Skeleton.Physics AdditionalPhysicsUpdate { get { return additionalPhysicsUpdate; } set { additionalPhysicsUpdate = value; } }
 		#endregion
 
 		public override void Initialize (bool overwrite, bool quiet = false) {
@@ -156,11 +170,11 @@ namespace Spine.Unity {
 			if (_UpdateLocal != null)
 				_UpdateLocal(this);
 
-			UpdateWorldTransform();
+			UpdateWorldTransform(mainPhysicsUpdate);
 
 			if (_UpdateWorld != null) {
 				_UpdateWorld(this);
-				UpdateWorldTransform();
+				UpdateWorldTransform(additionalPhysicsUpdate);
 			}
 
 			if (_UpdateComplete != null)
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
index b2c2123cc..b2dcf7c1c 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
@@ -423,7 +423,7 @@ namespace Spine.Unity {
 			// Generate mesh once, required to update mesh bounds for visibility
 			UpdateMode updateModeSaved = updateMode;
 			updateMode = UpdateMode.FullUpdate;
-			UpdateWorldTransform();
+			UpdateWorldTransform(Skeleton.Physics.Update);
 			LateUpdate();
 			updateMode = updateModeSaved;
 
@@ -439,8 +439,8 @@ namespace Spine.Unity {
 #endif
 		}
 
-		protected virtual void UpdateWorldTransform () {
-			skeleton.UpdateWorldTransform(Skeleton.Physics.Update);
+		protected virtual void UpdateWorldTransform (Skeleton.Physics physics) {
+			skeleton.UpdateWorldTransform(physics);
 		}
 
 		/// <summary>
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/ISkeletonAnimation.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/ISkeletonAnimation.cs
index 06dea7f70..67871d778 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/ISkeletonAnimation.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/ISkeletonAnimation.cs
@@ -63,6 +63,8 @@ namespace Spine.Unity {
 		event UpdateBonesDelegate UpdateComplete;
 		Skeleton Skeleton { get; }
 		UpdateTiming UpdateTiming { get; set; }
+		Skeleton.Physics MainPhysicsUpdate { get; set; }
+		Skeleton.Physics AdditionalPhysicsUpdate { get; set; }
 	}
 
 	/// <summary>Holds a reference to a SkeletonDataAsset.</summary>
diff --git a/spine-unity/Assets/Spine/package.json b/spine-unity/Assets/Spine/package.json
index 009ffc207..2e975374b 100644
--- a/spine-unity/Assets/Spine/package.json
+++ b/spine-unity/Assets/Spine/package.json
@@ -2,7 +2,7 @@
 	"name": "com.esotericsoftware.spine.spine-unity",
 	"displayName": "spine-unity Runtime",
 	"description": "This plugin provides the spine-unity runtime core.",
-	"version": "4.2.35",
+	"version": "4.2.36",
 	"unity": "2018.3",
 	"author": {
 		"name": "Esoteric Software",