diff --git a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Common-URP.hlsl b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Common-URP.hlsl
index f338e29c1..d40855e2c 100644
--- a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Common-URP.hlsl
+++ b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Common-URP.hlsl
@@ -1,6 +1,16 @@
 #ifndef SPINE_COMMON_URP_INCLUDED
 #define SPINE_COMMON_URP_INCLUDED
 
+#ifdef USE_FORWARD_PLUS
+#define IS_URP_14_OR_NEWER 1
+#else
+#define IS_URP_14_OR_NEWER 0
+#endif
+
+#if defined(_WRITE_RENDERING_LAYERS) && IS_URP_14_OR_NEWER
+#define USE_WRITE_RENDERING_LAYERS
+#endif
+
 #ifdef _LIGHT_LAYERS
 uint GetMeshRenderingLayerBackwardsCompatible()
 {
diff --git a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-SkeletonLit-ForwardPass-URP.hlsl b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-SkeletonLit-ForwardPass-URP.hlsl
index 98582e4ee..e40d392b2 100644
--- a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-SkeletonLit-ForwardPass-URP.hlsl
+++ b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-SkeletonLit-ForwardPass-URP.hlsl
@@ -152,7 +152,7 @@ VertexOutput vert(appdata v) {
 }
 
 half4 frag(VertexOutput i
-#ifdef _WRITE_RENDERING_LAYERS
+#ifdef USE_WRITE_RENDERING_LAYERS
 	, out float4 outRenderingLayers : SV_Target1
 #endif
 ) : SV_Target0
diff --git a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Sprite-ForwardPass-URP.hlsl b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Sprite-ForwardPass-URP.hlsl
index 146be9be8..303fcd1fd 100644
--- a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Sprite-ForwardPass-URP.hlsl
+++ b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/Shaders/Include/Spine-Sprite-ForwardPass-URP.hlsl
@@ -285,7 +285,7 @@ VertexOutputLWRP ForwardPassVertexSprite(VertexInput input)
 }
 
 half4 ForwardPassFragmentSprite(VertexOutputLWRP input
-#ifdef _WRITE_RENDERING_LAYERS
+#ifdef USE_WRITE_RENDERING_LAYERS
 	, out float4 outRenderingLayers : SV_Target1
 #endif
 ) : SV_Target0
@@ -302,7 +302,7 @@ half4 ForwardPassFragmentSprite(VertexOutputLWRP input
 	#if defined(REQUIRES_VERTEX_SHADOW_COORD_INTERPOLATOR)
 		inputData.shadowCoord = input.shadowCoord;
 	#elif defined(MAIN_LIGHT_CALCULATE_SHADOWS)
-		inputData.shadowCoord = TransformWorldToShadowCoord(input.positionWS);
+		inputData.shadowCoord = TransformWorldToShadowCoord(input.positionWS.xyz);
 	#elif defined(_MAIN_LIGHT_SHADOWS)
 		inputData.shadowCoord = input.shadowCoord;
 	#else
@@ -352,7 +352,7 @@ half4 ForwardPassFragmentSprite(VertexOutputLWRP input
 	COLORISE(pixel)
 	APPLY_FOG_LWRP(pixel, input.fogFactorAndVertexLight.x)
 
-#ifdef _WRITE_RENDERING_LAYERS
+#ifdef USE_WRITE_RENDERING_LAYERS
 	uint renderingLayers = GetMeshRenderingLayerBackwardsCompatible();
 	outRenderingLayers = float4(EncodeMeshRenderingLayer(renderingLayers), 0, 0, 0);
 #endif
diff --git a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/package.json b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/package.json
index 38dc2f4e2..b9cedf465 100644
--- a/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/package.json
+++ b/spine-unity/Modules/com.esotericsoftware.spine.urp-shaders/package.json
@@ -2,7 +2,7 @@
   "name": "com.esotericsoftware.spine.urp-shaders",
   "displayName": "Spine Universal RP Shaders",
   "description": "This plugin provides universal render pipeline (URP) shaders for the spine-unity runtime.\n\nPrerequisites:\nIt requires a working installation of the spine-unity runtime, version 4.1.\n(See http://esotericsoftware.com/git/spine-runtimes/spine-unity)",
-  "version": "4.1.9",
+  "version": "4.1.10",
   "unity": "2019.3",
   "author": {
     "name": "Esoteric Software",