diff --git a/spine-c/spine-c/src/spine/IkConstraint.c b/spine-c/spine-c/src/spine/IkConstraint.c
index 7ce325d8c..4f0fa257a 100644
--- a/spine-c/spine-c/src/spine/IkConstraint.c
+++ b/spine-c/spine-c/src/spine/IkConstraint.c
@@ -80,8 +80,8 @@ void spIkConstraint_apply1(spBone *bone, float targetX, float targetY, int /*boo
 
 	switch (bone->data->transformMode) {
 		case SP_TRANSFORMMODE_ONLYTRANSLATION:
-			tx = targetX - bone->worldX;
-			ty = targetY - bone->worldY;
+			tx = (targetX - bone->worldX) * SIGNUM(bone->skeleton->scaleX);
+			ty = (targetY - bone->worldY) * SIGNUM(bone->skeleton->scaleY);
 			break;
 		case SP_TRANSFORMMODE_NOROTATIONORREFLECTION: {
 			s = ABS(pa * pd - pb * pc) / MAX(0.0001f, pa * pa + pc * pc);
diff --git a/spine-cpp/spine-cpp/src/spine/IkConstraint.cpp b/spine-cpp/spine-cpp/src/spine/IkConstraint.cpp
index ac865f111..bf667d6b7 100644
--- a/spine-cpp/spine-cpp/src/spine/IkConstraint.cpp
+++ b/spine-cpp/spine-cpp/src/spine/IkConstraint.cpp
@@ -47,8 +47,8 @@ void IkConstraint::apply(Bone &bone, float targetX, float targetY, bool compress
 
 	switch (bone._data.getTransformMode()) {
 		case TransformMode_OnlyTranslation:
-			tx = targetX - bone._worldX;
-			ty = targetY - bone._worldY;
+			tx = (targetX - bone._worldX) * MathUtil::sign(bone.getSkeleton().getScaleX());
+			ty = (targetY - bone._worldY) * MathUtil::sign(bone.getSkeleton().getScaleY());
 			break;
 		case TransformMode_NoRotationOrReflection: {
 			float s = MathUtil::abs(pa * pd - pb * pc) / MathUtil::max(0.0001f, pa * pa + pc * pc);
diff --git a/spine-flutter/CHANGELOG.md b/spine-flutter/CHANGELOG.md
index c962b75c9..0929f26fb 100644
--- a/spine-flutter/CHANGELOG.md
+++ b/spine-flutter/CHANGELOG.md
@@ -1,5 +1,10 @@
 # 4.2.18
 * Fixes compilation errors due to API change in Flutter 3.16.0, see [this issue](https://github.com/EsotericSoftware/spine-runtimes/issues/2420). **Note**: Depending on this version requires your project to depend on Flutter >= 3.16.0 as well.
+# 4.1.10
+# Update WASM binaries
+
+# 4.1.9
+# Fixes #2412, single bone, translation only IK constraints did not take skeleton scale into account.
 
 # 4.1.8
 * Fixes compilation errors due to API change in Flutter 3.16.0, see [this issue](https://github.com/EsotericSoftware/spine-runtimes/issues/2420). **Note**: Depending on this version requires your project to depend on Flutter >= 3.16.0 as well.
diff --git a/spine-haxe/spine-haxe/spine/IkConstraint.hx b/spine-haxe/spine-haxe/spine/IkConstraint.hx
index 937d0a416..1e86eea00 100644
--- a/spine-haxe/spine-haxe/spine/IkConstraint.hx
+++ b/spine-haxe/spine-haxe/spine/IkConstraint.hx
@@ -95,8 +95,8 @@ class IkConstraint implements Updatable {
 			ty:Float = 0;
 		switch (bone.data.transformMode) {
 			case TransformMode.onlyTranslation:
-				tx = targetX - bone.worldX;
-				ty = targetY - bone.worldY;
+				tx = (targetX - bone.worldX) * MathUtils.signum(bone.skeleton.scaleX);
+				ty = (targetY - bone.worldY) * MathUtils.signum(bone.skeleton.scaleY);
 			case TransformMode.noRotationOrReflection:
 				var s = Math.abs(pa * pd - pb * pc) / Math.max(0.0001, pa * pa + pc * pc);
 				var sa:Float = pa / bone.skeleton.scaleX;
diff --git a/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/IkConstraint.java b/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/IkConstraint.java
index ef8192f08..c4d6be61f 100644
--- a/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/IkConstraint.java
+++ b/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/IkConstraint.java
@@ -191,8 +191,8 @@ public class IkConstraint implements Updatable {
 		float rotationIK = -bone.ashearX - bone.arotation, tx, ty;
 		switch (bone.data.transformMode) {
 		case onlyTranslation:
-			tx = targetX - bone.worldX;
-			ty = targetY - bone.worldY;
+			tx = (targetX - bone.worldX) * Math.signum(bone.skeleton.scaleX);
+			ty = (targetY - bone.worldY) * Math.signum(bone.skeleton.scaleY);
 			break;
 		case noRotationOrReflection:
 			float s = Math.abs(pa * pd - pb * pc) / Math.max(0.0001f, pa * pa + pc * pc);
diff --git a/spine-ts/spine-core/src/IkConstraint.ts b/spine-ts/spine-core/src/IkConstraint.ts
index de5045a63..9fc98ff63 100644
--- a/spine-ts/spine-core/src/IkConstraint.ts
+++ b/spine-ts/spine-core/src/IkConstraint.ts
@@ -122,8 +122,8 @@ export class IkConstraint implements Updatable {
 
 		switch (bone.data.transformMode) {
 			case TransformMode.OnlyTranslation:
-				tx = targetX - bone.worldX;
-				ty = targetY - bone.worldY;
+				tx = (targetX - bone.worldX) * MathUtils.signum(bone.skeleton.scaleX);
+				ty = (targetY - bone.worldY) * MathUtils.signum(bone.skeleton.scaleY);
 				break;
 			case TransformMode.NoRotationOrReflection:
 				let s = Math.abs(pa * pd - pb * pc) / Math.max(0.0001, pa * pa + pc * pc);