diff --git a/spine-cpp/spine-cpp/include/spine/BoneLocal.h b/spine-cpp/spine-cpp/include/spine/BoneLocal.h
index 35f0a6d01..e9969eeab 100644
--- a/spine-cpp/spine-cpp/include/spine/BoneLocal.h
+++ b/spine-cpp/spine-cpp/include/spine/BoneLocal.h
@@ -38,6 +38,9 @@
 namespace spine {
 	/// Stores a bone's local pose.
 	class SP_API BoneLocal : public Pose<BoneLocal> {
+		friend class BoneTimeline1;
+		friend class RotateTimeline;
+
 		RTTI_DECL
 
 	protected:
diff --git a/spine-cpp/spine-cpp/include/spine/BoneTimeline.h b/spine-cpp/spine-cpp/include/spine/BoneTimeline.h
index 9ea945c01..cc1664469 100644
--- a/spine-cpp/spine-cpp/include/spine/BoneTimeline.h
+++ b/spine-cpp/spine-cpp/include/spine/BoneTimeline.h
@@ -31,8 +31,13 @@
 #define Spine_BoneTimeline_h
 
 #include <spine/dll.h>
+#include <spine/CurveTimeline.h>
 
 namespace spine {
+	class Skeleton;
+	class Event;
+	class BoneLocal;
+
     /// An interface for timelines which change the property of a bone.
     class SP_API BoneTimeline {
     public:
@@ -42,6 +47,33 @@ namespace spine {
         /// The index of the bone in Skeleton::getBones() that will be changed when this timeline is applied.
         virtual int getBoneIndex() = 0;
     };
+
+	/// Base class for timelines that animate a single bone property.
+	class SP_API BoneTimeline1 : public CurveTimeline1, public BoneTimeline {
+		friend class SkeletonBinary;
+		friend class SkeletonJson;
+		friend class AnimationState;
+
+	RTTI_DECL
+
+	public:
+		BoneTimeline1(size_t frameCount, size_t bezierCount, int boneIndex, Property property);
+
+		virtual void
+		apply(Skeleton &skeleton, float lastTime, float time, Vector<Event *> *pEvents, float alpha, MixBlend blend,
+			  MixDirection direction, bool appliedPose) override;
+
+		virtual int getBoneIndex() override { return _boneIndex; }
+
+		void setBoneIndex(int inValue) { _boneIndex = inValue; }
+
+	protected:
+		/// Applies changes to the pose based on the timeline values.
+		virtual void apply(BoneLocal &pose, BoneLocal &setup, float time, float alpha, MixBlend blend,
+						   MixDirection direction) = 0;
+
+		int _boneIndex;
+	};
 }
 
 #endif
\ No newline at end of file
diff --git a/spine-cpp/spine-cpp/include/spine/Posed.h b/spine-cpp/spine-cpp/include/spine/Posed.h
index 1f1d5ffbf..ea298421f 100644
--- a/spine-cpp/spine-cpp/include/spine/Posed.h
+++ b/spine-cpp/spine-cpp/include/spine/Posed.h
@@ -37,6 +37,7 @@ namespace spine {
 	template<class D, class P, class A>
 	class SP_API Posed : public SpineObject {
 		friend class AnimationState;
+		friend class BoneTimeline1;
 		friend class RotateTimeline;
 		friend class IkConstraint;
 		friend class TransformConstraint;
diff --git a/spine-cpp/spine-cpp/include/spine/PosedData.h b/spine-cpp/spine-cpp/include/spine/PosedData.h
index b3aa8ad4b..b8f07a4e9 100644
--- a/spine-cpp/spine-cpp/include/spine/PosedData.h
+++ b/spine-cpp/spine-cpp/include/spine/PosedData.h
@@ -42,6 +42,24 @@ namespace spine {
     class SP_API PosedData : public SpineObject {
         friend class SkeletonBinary;
         friend class SkeletonJson;
+        friend class BoneTimeline1;
+        friend class RotateTimeline;
+        friend class AttachmentTimeline;
+        friend class RGBATimeline;
+        friend class RGBTimeline;
+        friend class AlphaTimeline;
+        friend class RGBA2Timeline;
+        friend class RGB2Timeline;
+        friend class ScaleTimeline;
+        friend class ScaleXTimeline;
+        friend class ScaleYTimeline;
+        friend class ShearTimeline;
+        friend class ShearXTimeline;
+        friend class ShearYTimeline;
+        friend class TranslateTimeline;
+        friend class TranslateXTimeline;
+        friend class TranslateYTimeline;
+        friend class InheritTimeline;
 
     private:
         spine::String _name;
diff --git a/spine-cpp/spine-cpp/include/spine/RotateTimeline.h b/spine-cpp/spine-cpp/include/spine/RotateTimeline.h
index 89b8621f9..086217fb3 100644
--- a/spine-cpp/spine-cpp/include/spine/RotateTimeline.h
+++ b/spine-cpp/spine-cpp/include/spine/RotateTimeline.h
@@ -30,10 +30,11 @@
 #ifndef Spine_RotateTimeline_h
 #define Spine_RotateTimeline_h
 
-#include <spine/CurveTimeline.h>
+#include <spine/BoneTimeline.h>
 
 namespace spine {
-	class SP_API RotateTimeline : public CurveTimeline1 {
+	/// Changes a bone's local rotation.
+	class SP_API RotateTimeline : public BoneTimeline1 {
 		friend class SkeletonBinary;
 
 		friend class SkeletonJson;
@@ -45,16 +46,9 @@ namespace spine {
 	public:
 		explicit RotateTimeline(size_t frameCount, size_t bezierCount, int boneIndex);
 
-		virtual void
-		apply(Skeleton &skeleton, float lastTime, float time, Vector<Event *> *pEvents, float alpha, MixBlend blend,
-			  MixDirection direction);
-
-		int getBoneIndex() { return _boneIndex; }
-
-		void setBoneIndex(int inValue) { _boneIndex = inValue; }
-
-	private:
-		int _boneIndex;
+	protected:
+		virtual void apply(BoneLocal &pose, BoneLocal &setup, float time, float alpha, MixBlend blend,
+						   MixDirection direction) override;
 	};
 }
 
diff --git a/spine-cpp/spine-cpp/include/spine/Skeleton.h b/spine-cpp/spine-cpp/include/spine/Skeleton.h
index 82304d6a9..cb97aba92 100644
--- a/spine-cpp/spine-cpp/include/spine/Skeleton.h
+++ b/spine-cpp/spine-cpp/include/spine/Skeleton.h
@@ -107,6 +107,8 @@ namespace spine {
 
 		friend class TransformConstraintTimeline;
 
+		friend class BoneTimeline1;
+
 		friend class RotateTimeline;
 
 		friend class TranslateTimeline;
diff --git a/spine-cpp/spine-cpp/src/spine/BoneTimeline.cpp b/spine-cpp/spine-cpp/src/spine/BoneTimeline.cpp
index fae0dd334..a56ee4023 100644
--- a/spine-cpp/spine-cpp/src/spine/BoneTimeline.cpp
+++ b/spine-cpp/spine-cpp/src/spine/BoneTimeline.cpp
@@ -29,10 +29,36 @@
 
 #include <spine/BoneTimeline.h>
 
+#include <spine/Event.h>
+#include <spine/Skeleton.h>
+#include <spine/Bone.h>
+#include <spine/BoneData.h>
+#include <spine/BoneLocal.h>
+#include <spine/Property.h>
+
 using namespace spine;
 
 BoneTimeline::BoneTimeline() {
 }
 
 BoneTimeline::~BoneTimeline() {
+}
+
+RTTI_IMPL(BoneTimeline1, CurveTimeline1)
+
+BoneTimeline1::BoneTimeline1(size_t frameCount, size_t bezierCount, int boneIndex, Property property) : 
+	CurveTimeline1(frameCount, bezierCount), _boneIndex(boneIndex) {
+	PropertyId ids[] = {((PropertyId) property << 32) | boneIndex};
+	setPropertyIds(ids, 1);
+}
+
+void BoneTimeline1::apply(Skeleton &skeleton, float lastTime, float time, Vector<Event *> *pEvents, float alpha,
+						  MixBlend blend, MixDirection direction, bool appliedPose) {
+	SP_UNUSED(lastTime);
+	SP_UNUSED(pEvents);
+
+	Bone *bone = skeleton._bones[_boneIndex];
+	if (bone->isActive()) {
+		apply(appliedPose ? *bone->_applied : bone->_pose, *bone->_data._setup, time, alpha, blend, direction);
+	}
 }
\ No newline at end of file
diff --git a/spine-cpp/spine-cpp/src/spine/RotateTimeline.cpp b/spine-cpp/spine-cpp/src/spine/RotateTimeline.cpp
index 4c53f3010..50ef5d8b6 100644
--- a/spine-cpp/spine-cpp/src/spine/RotateTimeline.cpp
+++ b/spine-cpp/spine-cpp/src/spine/RotateTimeline.cpp
@@ -29,31 +29,18 @@
 
 #include <spine/RotateTimeline.h>
 
-#include <spine/Event.h>
-#include <spine/Skeleton.h>
-
-#include <spine/Animation.h>
-#include <spine/Bone.h>
-#include <spine/BoneData.h>
-#include <spine/Property.h>
+#include <spine/BoneLocal.h>
 
 using namespace spine;
 
-RTTI_IMPL(RotateTimeline, CurveTimeline1)
+RTTI_IMPL(RotateTimeline, BoneTimeline1)
 
-RotateTimeline::RotateTimeline(size_t frameCount, size_t bezierCount, int boneIndex) : CurveTimeline1(frameCount,
-																									  bezierCount),
-																					   _boneIndex(boneIndex) {
-	PropertyId ids[] = {((PropertyId) Property_Rotate << 32) | boneIndex};
-	setPropertyIds(ids, 1);
+RotateTimeline::RotateTimeline(size_t frameCount, size_t bezierCount, int boneIndex) : 
+	BoneTimeline1(frameCount, bezierCount, boneIndex, Property_Rotate) {
 }
 
-void RotateTimeline::apply(Skeleton &skeleton, float lastTime, float time, Vector<Event *> *pEvents, float alpha,
-						   MixBlend blend, MixDirection direction) {
-	SP_UNUSED(lastTime);
-	SP_UNUSED(pEvents);
+void RotateTimeline::apply(BoneLocal &pose, BoneLocal &setup, float time, float alpha, MixBlend blend,
+						   MixDirection direction) {
 	SP_UNUSED(direction);
-
-	Bone *bone = skeleton._bones[_boneIndex];
-	if (bone->isActive()) bone->_rotation = getRelativeValue(time, alpha, blend, bone->_rotation, bone->getData()._rotation);
-}
+	pose._rotation = getRelativeValue(time, alpha, blend, pose._rotation, setup._rotation);
+}
\ No newline at end of file