diff --git a/spine-unity/Modules/com.esotericsoftware.spine.timeline/Runtime/SpineAnimationState/SpineAnimationStateMixerBehaviour.cs b/spine-unity/Modules/com.esotericsoftware.spine.timeline/Runtime/SpineAnimationState/SpineAnimationStateMixerBehaviour.cs
index a9e472020..bf25ef008 100644
--- a/spine-unity/Modules/com.esotericsoftware.spine.timeline/Runtime/SpineAnimationState/SpineAnimationStateMixerBehaviour.cs
+++ b/spine-unity/Modules/com.esotericsoftware.spine.timeline/Runtime/SpineAnimationState/SpineAnimationStateMixerBehaviour.cs
@@ -303,12 +303,14 @@ namespace Spine.Unity.Playables {
 					if (toAnimation != null)
 						toAnimation.Apply(skeleton, 0, toClipTime, clipData.loop, null, 1f, MixBlend.Setup, MixDirection.In);
 				}
-				skeleton.UpdateWorldTransform();
 
-				if (skeletonAnimation)
+				if (skeletonAnimation) {
+					skeletonAnimation.Update(0);
 					skeletonAnimation.LateUpdate();
-				else if (skeletonGraphic)
+				} else if (skeletonGraphic) {
+					skeletonGraphic.Update(0);
 					skeletonGraphic.LateUpdate();
+				}
 			}
 			// Do nothing outside of the first clip and the last clip.