diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
index b8268e5b4..d1f3cf5d3 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonAnimation.cs
@@ -244,12 +244,19 @@ namespace Spine.Unity {
 			skeleton.Update(deltaTime);
 
 			if (Application.isPlaying) {
-				Vector2 position = new Vector2(transform.position.x, transform.position.y);
-				Vector2 positionDelta = position - lastPosition;
-				positionDelta.x /= transform.lossyScale.x;
-				positionDelta.y /= transform.lossyScale.y;
-				skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
-				lastPosition = position;
+				if (applyTranslationToPhysics) {
+					Vector2 position = new Vector2(transform.position.x, transform.position.y);
+					Vector2 positionDelta = position - lastPosition;
+					positionDelta.x /= transform.lossyScale.x;
+					positionDelta.y /= transform.lossyScale.y;
+					skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
+					lastPosition = position;
+				}
+				if (applyRotationToPhysics) {
+					float rotation = this.transform.rotation.eulerAngles.z;
+					skeleton.PhysicsRotate(0, 0, rotation - lastRotation);
+					lastRotation = rotation;
+				}
 			}
 
 			if (updateMode == UpdateMode.OnlyAnimationStatus) {
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
index 826e4d62a..1c7ff9d38 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonGraphic.cs
@@ -382,12 +382,19 @@ namespace Spine.Unity {
 			skeleton.Update(deltaTime);
 
 			if (Application.isPlaying) {
-				Vector2 position = new Vector2(transform.position.x, transform.position.y);
-				Vector2 positionDelta = (position - lastPosition) / meshScale;
-				positionDelta.x /= transform.lossyScale.x;
-				positionDelta.y /= transform.lossyScale.y;
-				skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
-				lastPosition = position;
+				if (applyTranslationToPhysics) {
+					Vector2 position = new Vector2(transform.position.x, transform.position.y);
+					Vector2 positionDelta = (position - lastPosition) / meshScale;
+					positionDelta.x /= transform.lossyScale.x;
+					positionDelta.y /= transform.lossyScale.y;
+					skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
+					lastPosition = position;
+				}
+				if (applyRotationToPhysics) {
+					float rotation = this.transform.rotation.eulerAngles.z;
+					skeleton.PhysicsRotate(0, 0, rotation - lastRotation);
+					lastRotation = rotation;
+				}
 			}
 
 			if (updateMode == UpdateMode.OnlyAnimationStatus) {
@@ -528,8 +535,22 @@ namespace Spine.Unity {
 			}
 		}
 
-		/// <summary>Used for applying Transform translation to skeleton physics.</summary>
+		/// <summary>When enabled, Transform translation is applied to skeleton PhysicsConstraints.</summary>
+		public bool applyTranslationToPhysics = true;
+		/// <summary>When enabled, Transform rotation is applied to skeleton PhysicsConstraints.</summary>
+		public bool applyRotationToPhysics = true;
+
+		/// <summary>Used for applying Transform translation to skeleton PhysicsConstraints.</summary>
 		protected Vector2 lastPosition;
+		/// <summary>Used for applying Transform rotation to skeleton PhysicsConstraints.</summary>
+		protected float lastRotation;
+
+		public void ResetLastPosition () { lastPosition = this.transform.position; }
+		public void ResetLastRotation () { lastRotation = this.transform.rotation.eulerAngles.z; }
+		public void ResetLastPositionAndRotation () {
+			lastPosition = this.transform.position;
+			lastRotation = this.transform.rotation.eulerAngles.z;
+		}
 
 		[SerializeField] protected Spine.Unity.MeshGenerator meshGenerator = new MeshGenerator();
 		public Spine.Unity.MeshGenerator MeshGenerator { get { return this.meshGenerator; } }
@@ -690,6 +711,8 @@ namespace Spine.Unity {
 		}
 
 		public void Initialize (bool overwrite) {
+			ResetLastPositionAndRotation();
+
 			if (this.IsValid && !overwrite) return;
 #if UNITY_EDITOR
 			if (BuildUtilities.IsInSkeletonAssetBuildPreProcessing)
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
index ac87f089f..219dbf80e 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonMecanim.cs
@@ -127,12 +127,19 @@ namespace Spine.Unity {
 			skeleton.Update(deltaTime);
 
 			if (Application.isPlaying) {
-				Vector2 position = new Vector2(transform.position.x, transform.position.y);
-				Vector2 positionDelta = position - lastPosition;
-				positionDelta.x /= transform.lossyScale.x;
-				positionDelta.y /= transform.lossyScale.y;
-				skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
-				lastPosition = position;
+				if (applyTranslationToPhysics) {
+					Vector2 position = new Vector2(transform.position.x, transform.position.y);
+					Vector2 positionDelta = position - lastPosition;
+					positionDelta.x /= transform.lossyScale.x;
+					positionDelta.y /= transform.lossyScale.y;
+					skeleton.PhysicsTranslate(positionDelta.x, positionDelta.y);
+					lastPosition = position;
+				}
+				if (applyRotationToPhysics) {
+					float rotation = this.transform.rotation.eulerAngles.z;
+					skeleton.PhysicsRotate(0, 0, rotation - lastRotation);
+					lastRotation = rotation;
+				}
 			}
 
 			ApplyAnimation();
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
index dd05b9438..f85ca5e3e 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonRenderer.cs
@@ -286,9 +286,25 @@ namespace Spine.Unity {
 				return skeleton;
 			}
 		}
+		#endregion
 
-		/// <summary>Used for applying Transform translation to skeleton physics.</summary>
+		#region Physics
+		/// <summary>When enabled, Transform translation is applied to skeleton PhysicsConstraints.</summary>
+		public bool applyTranslationToPhysics = true;
+		/// <summary>When enabled, Transform rotation is applied to skeleton PhysicsConstraints.</summary>
+		public bool applyRotationToPhysics = true;
+
+		/// <summary>Used for applying Transform translation to skeleton PhysicsConstraints.</summary>
 		protected Vector2 lastPosition;
+		/// <summary>Used for applying Transform rotation to skeleton PhysicsConstraints.</summary>
+		protected float lastRotation;
+
+		public void ResetLastPosition () { lastPosition = this.transform.position; }
+		public void ResetLastRotation () { lastRotation = this.transform.rotation.eulerAngles.z; }
+		public void ResetLastPositionAndRotation () {
+			lastPosition = this.transform.position;
+			lastRotation = this.transform.rotation.eulerAngles.z;
+		}
 		#endregion
 
 		public delegate void SkeletonRendererDelegate (SkeletonRenderer skeletonRenderer);
@@ -380,6 +396,8 @@ namespace Spine.Unity {
 		/// Initialize this component. Attempts to load the SkeletonData and creates the internal Skeleton object and buffers.</summary>
 		/// <param name="overwrite">If set to <c>true</c>, it will overwrite internal objects if they were already generated. Otherwise, the initialized component will ignore subsequent calls to initialize.</param>
 		public virtual void Initialize (bool overwrite, bool quiet = false) {
+			ResetLastPositionAndRotation();
+
 			if (valid && !overwrite)
 				return;
 #if UNITY_EDITOR
@@ -427,7 +445,6 @@ namespace Spine.Unity {
 			UpdateMode updateModeSaved = updateMode;
 			updateMode = UpdateMode.FullUpdate;
 			UpdateWorldTransform(Skeleton.Physics.Update);
-			lastPosition = this.transform.position;
 			LateUpdate();
 			updateMode = updateModeSaved;
 
diff --git a/spine-unity/Assets/Spine/package.json b/spine-unity/Assets/Spine/package.json
index 12f964eae..0c4ebeddf 100644
--- a/spine-unity/Assets/Spine/package.json
+++ b/spine-unity/Assets/Spine/package.json
@@ -2,7 +2,7 @@
 	"name": "com.esotericsoftware.spine.spine-unity",
 	"displayName": "spine-unity Runtime",
 	"description": "This plugin provides the spine-unity runtime core.",
-	"version": "4.2.43",
+	"version": "4.2.44",
 	"unity": "2018.3",
 	"author": {
 		"name": "Esoteric Software",