diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtility.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtility.cs
index 1960bd3ef..76060edae 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtility.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtility.cs
@@ -204,8 +204,8 @@ namespace Spine.Unity {
 				UpdateLocal(skeletonGraphic);
 				UpdateWorld(skeletonGraphic);
 				UpdateComplete(skeletonGraphic);
- 			}
- 		}
+			}
+		}
 
 		[HideInInspector] public SkeletonRenderer skeletonRenderer;
 		[HideInInspector] public SkeletonGraphic skeletonGraphic;
@@ -362,16 +362,21 @@ namespace Spine.Unity {
 			if (skeleton == null) return;
 
 			if (boneRoot != null) {
-				List<object> constraintTargets = new List<object>();
+				List<Bone> constrainedBones = new List<Bone>();
 				ExposedList<IConstraint> constraints = skeleton.Constraints;
 				for (int i = 0, n = constraints.Count; i < n; i++) {
 					IConstraint constraint = constraints.Items[i];
+					ExposedList<BonePose> bones = null;
 					if (constraint is IkConstraint)
-						constraintTargets.Add(((IkConstraint)constraint).Bones);
+						bones = ((IkConstraint)constraint).Bones;
 					else if (constraint is TransformConstraint)
-						constraintTargets.Add(((TransformConstraint)constraint).Bones);
+						bones = ((TransformConstraint)constraint).Bones;
 					else if (constraint is PathConstraint)
-						constraintTargets.Add(((PathConstraint)constraint).Bones);
+						bones = ((PathConstraint)constraint).Bones;
+					if (bones != null) {
+						for (int j = 0, m = bones.Count; j < m; j++)
+							constrainedBones.Add(bones.Items[j].bone);
+					}
 				}
 
 				List<SkeletonUtilityBone> boneComponents = this.boneComponents;
@@ -382,7 +387,7 @@ namespace Spine.Unity {
 						if (b.bone == null) continue;
 					}
 					hasOverrideBones |= (b.mode == SkeletonUtilityBone.Mode.Override);
-					hasConstraintTargetBones |= constraintTargets.Contains(b.bone);
+					hasConstraintTargetBones |= constrainedBones.Contains(b.bone);
 				}
 
 				needToReprocessBones = false;
diff --git a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtilityBone.cs b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtilityBone.cs
index 39681a07f..f37d9f457 100644
--- a/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtilityBone.cs
+++ b/spine-unity/Assets/Spine/Runtime/spine-unity/Components/SkeletonUtility/SkeletonUtilityBone.cs
@@ -54,7 +54,7 @@ namespace Spine.Unity {
 			Complete
 		}
 
-#region Inspector
+		#region Inspector
 		/// <summary>If a bone isn't set, boneName is used to find the bone.</summary>
 		public string boneName;
 		public Transform parentReference;
@@ -62,7 +62,7 @@ namespace Spine.Unity {
 		public bool position, rotation, scale, zPosition = true;
 		[Range(0f, 1f)]
 		public float overrideAlpha = 1;
-#endregion
+		#endregion
 
 		public SkeletonUtility hierarchy;
 		[System.NonSerialized] public Bone bone;
@@ -159,6 +159,7 @@ namespace Spine.Unity {
 				case UpdatePhase.World:
 				case UpdatePhase.Complete:
 					var appliedPose = bone.AppliedPose;
+					appliedPose.ValidateLocalTransform(skeleton);
 					if (position)
 						thisTransform.localPosition = new Vector3(appliedPose.X * positionScale, appliedPose.Y * positionScale,
 							zPosition ? 0 : thisTransform.localPosition.z);
diff --git a/spine-unity/Assets/Spine/package.json b/spine-unity/Assets/Spine/package.json
index 94861664d..544e28858 100644
--- a/spine-unity/Assets/Spine/package.json
+++ b/spine-unity/Assets/Spine/package.json
@@ -2,7 +2,7 @@
 	"name": "com.esotericsoftware.spine.spine-unity",
 	"displayName": "spine-unity Runtime",
 	"description": "This plugin provides the spine-unity runtime core and examples. Spine Examples can be installed via the Samples tab.",
-	"version": "4.3.48",
+	"version": "4.3.49",
 	"unity": "2018.3",
 	"author": {
 		"name": "Esoteric Software",