diff --git a/spine-c/spine-c/src/spine/AnimationState.c b/spine-c/spine-c/src/spine/AnimationState.c
index e6ede4220..2ff7f484c 100644
--- a/spine-c/spine-c/src/spine/AnimationState.c
+++ b/spine-c/spine-c/src/spine/AnimationState.c
@@ -595,7 +595,8 @@ _spAnimationState_setAttachment(spAnimationState *self, spSkeleton *skeleton, sp
 
 /* @param target After the first and before the last entry. */
 static int binarySearch1(float *values, int valuesLength, float target) {
-	for (int i = 1; i < valuesLength; i++) {
+	int i;
+	for (i = 1; i < valuesLength; i++) {
 		if (values[i] > target) return (int) (i - 1);
 	}
 	return (int) valuesLength - 1;
diff --git a/spine-c/spine-c/src/spine/RegionAttachment.c b/spine-c/spine-c/src/spine/RegionAttachment.c
index f888d6100..b91ce5bef 100644
--- a/spine-c/spine-c/src/spine/RegionAttachment.c
+++ b/spine-c/spine-c/src/spine/RegionAttachment.c
@@ -79,22 +79,39 @@ spRegionAttachment *spRegionAttachment_create(const char *name) {
 }
 
 void spRegionAttachment_updateRegion(spRegionAttachment *self) {
-	float regionScaleX = self->width / self->region->originalWidth * self->scaleX;
-	float regionScaleY = self->height / self->region->originalHeight * self->scaleY;
-	float localX = -self->width / 2 * self->scaleX + self->region->offsetX * regionScaleX;
-	float localY = -self->height / 2 * self->scaleY + self->region->offsetY * regionScaleY;
-	float localX2 = localX + self->region->width * regionScaleX;
-	float localY2 = localY + self->region->height * regionScaleY;
-	float radians = self->rotation * DEG_RAD;
-	float cosine = COS(radians), sine = SIN(radians);
-	float localXCos = localX * cosine + self->x;
-	float localXSin = localX * sine;
-	float localYCos = localY * cosine + self->y;
-	float localYSin = localY * sine;
-	float localX2Cos = localX2 * cosine + self->x;
-	float localX2Sin = localX2 * sine;
-	float localY2Cos = localY2 * cosine + self->y;
-	float localY2Sin = localY2 * sine;
+	float regionScaleX, regionScaleY, localX, localY, localX2, localY2;
+	float radians, cosine, sine;
+	float localXCos, localXSin, localYCos, localYSin, localX2Cos, localX2Sin, localY2Cos, localY2Sin;
+
+	if (self->region == NULL) {
+		self->uvs[0] = 0;
+		self->uvs[1] = 0;
+		self->uvs[2] = 1;
+		self->uvs[3] = 1;
+		self->uvs[4] = 1;
+		self->uvs[5] = 0;
+		self->uvs[6] = 0;
+		self->uvs[7] = 0;
+		return;
+	}
+
+	regionScaleX = self->width / self->region->originalWidth * self->scaleX;
+	regionScaleY = self->height / self->region->originalHeight * self->scaleY;
+	localX = -self->width / 2 * self->scaleX + self->region->offsetX * regionScaleX;
+	localY = -self->height / 2 * self->scaleY + self->region->offsetY * regionScaleY;
+	localX2 = localX + self->region->width * regionScaleX;
+	localY2 = localY + self->region->height * regionScaleY;
+	radians = self->rotation * DEG_RAD;
+	cosine = COS(radians), sine = SIN(radians);
+	localXCos = localX * cosine + self->x;
+	localXSin = localX * sine;
+	localYCos = localY * cosine + self->y;
+	localYSin = localY * sine;
+	localX2Cos = localX2 * cosine + self->x;
+	localX2Sin = localX2 * sine;
+	localY2Cos = localY2 * cosine + self->y;
+	localY2Sin = localY2 * sine;
+
 	self->offset[BLX] = localXCos - localYSin;
 	self->offset[BLY] = localYCos + localXSin;
 	self->offset[ULX] = localXCos - localY2Sin;
diff --git a/spine-c/spine-c/src/spine/Skeleton.c b/spine-c/spine-c/src/spine/Skeleton.c
index 953f36e86..a637ba6f8 100644
--- a/spine-c/spine-c/src/spine/Skeleton.c
+++ b/spine-c/spine-c/src/spine/Skeleton.c
@@ -437,7 +437,6 @@ void spSkeleton_updateWorldTransformWith(const spSkeleton *self, const spBone *p
 	/* Apply the parent bone transform to the root bone. The root bone always inherits scale, rotation and reflection. */
 	int i;
 	float rotationY, la, lb, lc, ld;
-	_spUpdate *updateCache;
 	_spSkeleton *internal = SUB_CAST(_spSkeleton, self);
 	spBone *rootBone = self->root;
 	float pa = parent->a, pb = parent->b, pc = parent->c, pd = parent->d;
@@ -455,7 +454,6 @@ void spSkeleton_updateWorldTransformWith(const spSkeleton *self, const spBone *p
 	CONST_CAST(float, rootBone->d) = (pc * lb + pd * ld) * self->scaleY;
 
 	/* Update everything except root bone. */
-	updateCache = internal->updateCache;
 	for (i = 0; i < internal->updateCacheCount; ++i) {
 		_spUpdate *update = internal->updateCache + i;
 		switch (update->type) {
diff --git a/spine-cocos2dx/README.md b/spine-cocos2dx/README.md
index e169faae9..a025533b8 100644
--- a/spine-cocos2dx/README.md
+++ b/spine-cocos2dx/README.md
@@ -68,13 +68,13 @@ Execute the following on the command line:
 
 ```
 cd spine-runtimes/spine-cocos2dx/example
-mkdir build-macos && cmake . -GXcode -Bbuild-macos
+mkdir build-macos && cmake . -GXcode -Bbuild-macos -DCMAKE_OSX_ARCHITECTURES=x86_64
 open build-macos/spine-cocos2dx-example.xcodeproj
 ```
 
 This will generate an Xcode project in `build-macos/spine-cocos2dx-example.xcodeproj` and open it in Xcode. To build and run the example, select the `spine-cocos2dx-example` scheme and press `CMD + R`.
 
-> **Note**: cocos2d-x only ships pre-built external libraries like Bullet or FreeType for x86_64. If you build on a machine with Apple Silicon, adjust the `cmake` command above to `mkdir build-macos && cmake . -GXcode -Bbuild-macos -DCMAKE_OSX_ARCHITECTURES=x86_64`.
+> **NOTE:** Passing `-DCMAKE_OSX_ARCHITECTURES=x86_64` to CMake is currently required, as Cocos2d-X only provides prebuilt x86_64 binaries for its external dependencies.
 
 ### iOS
 Execute the following on the command line:
diff --git a/spine-cpp/spine-cpp/src/spine/RegionAttachment.cpp b/spine-cpp/spine-cpp/src/spine/RegionAttachment.cpp
index da15c053c..a1024d193 100644
--- a/spine-cpp/spine-cpp/src/spine/RegionAttachment.cpp
+++ b/spine-cpp/spine-cpp/src/spine/RegionAttachment.cpp
@@ -68,6 +68,18 @@ RegionAttachment::~RegionAttachment() {
 }
 
 void RegionAttachment::updateRegion() {
+	if (_region == NULL) {
+		_uvs[BLX] = 0;
+		_uvs[BLY] = 0;
+		_uvs[ULX] = 0;
+		_uvs[ULY] = 1;
+		_uvs[URX] = 1;
+		_uvs[URY] = 1;
+		_uvs[BRX] = 1;
+		_uvs[BRY] = 0;
+		return;
+	}
+
 	float regionScaleX = _width / _region->originalWidth * _scaleX;
 	float regionScaleY = _height / _region->originalHeight * _scaleY;
 	float localX = -_width / 2 * _scaleX + _region->offsetX * regionScaleX;
diff --git a/spine-csharp/src/Attachments/RegionAttachment.cs b/spine-csharp/src/Attachments/RegionAttachment.cs
index 0e7197ea7..1e1ec929e 100644
--- a/spine-csharp/src/Attachments/RegionAttachment.cs
+++ b/spine-csharp/src/Attachments/RegionAttachment.cs
@@ -97,7 +97,7 @@ namespace Spine {
 			if (region == null) {
 				uvs[BLX] = 0;
 				uvs[BLY] = 0;
-				uvs[ULX] = 1;
+				uvs[ULX] = 0;
 				uvs[ULY] = 1;
 				uvs[URX] = 1;
 				uvs[URY] = 1;
diff --git a/spine-godot/spine_godot/SpineSprite.cpp b/spine-godot/spine_godot/SpineSprite.cpp
index 68d2c7b96..9cbc69923 100644
--- a/spine-godot/spine_godot/SpineSprite.cpp
+++ b/spine-godot/spine_godot/SpineSprite.cpp
@@ -552,10 +552,10 @@ void SpineSprite::update_meshes(Ref<SpineSkeleton> skeleton_ref) {
 
 		if (attachment->getRTTI().isExactly(spine::RegionAttachment::rtti)) {
 			auto *region = (spine::RegionAttachment *) attachment;
-			renderer_object = (SpineRendererObject *) region->getRegion()->rendererObject;
 
 			vertices->setSize(8, 0);
 			region->computeWorldVertices(*slot, *vertices, 0);
+			renderer_object = (SpineRendererObject *) ((spine::AtlasRegion *) region->getRendererObject())->page->getRendererObject();
 			uvs = &region->getUVs();
 			indices = &quad_indices;
 
@@ -566,10 +566,10 @@ void SpineSprite::update_meshes(Ref<SpineSkeleton> skeleton_ref) {
 			tint.a *= attachment_color.a;
 		} else if (attachment->getRTTI().isExactly(spine::MeshAttachment::rtti)) {
 			auto *mesh = (spine::MeshAttachment *) attachment;
-			renderer_object = (SpineRendererObject *) mesh->getRegion()->rendererObject;
 
 			vertices->setSize(mesh->getWorldVerticesLength(), 0);
 			mesh->computeWorldVertices(*slot, *vertices);
+			renderer_object = (SpineRendererObject *) ((spine::AtlasRegion *) mesh->getRendererObject())->page->getRendererObject();
 			uvs = &mesh->getUVs();
 			indices = &mesh->getTriangles();
 
diff --git a/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/attachments/RegionAttachment.java b/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/attachments/RegionAttachment.java
index 5b439df7d..2ea4d4a90 100644
--- a/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/attachments/RegionAttachment.java
+++ b/spine-libgdx/spine-libgdx/src/com/esotericsoftware/spine/attachments/RegionAttachment.java
@@ -84,7 +84,7 @@ public class RegionAttachment extends Attachment implements HasTextureRegion {
 		if (region == null) {
 			uvs[BLX] = 0;
 			uvs[BLY] = 0;
-			uvs[ULX] = 1;
+			uvs[ULX] = 0;
 			uvs[ULY] = 1;
 			uvs[URX] = 1;
 			uvs[URY] = 1;
diff --git a/spine-sdl/README.md b/spine-sdl/README.md
index cf91c35f1..f02be0700 100644
--- a/spine-sdl/README.md
+++ b/spine-sdl/README.md
@@ -26,7 +26,7 @@ spine-sdl supports all Spine features except premultiplied alpha, screen blend m
 1. Create a new SDL project. See the [SDL documentation](https://wiki.libsdl.org/FrontPage) or have a look at the example in this repository.
 2. Download the Spine Runtimes source using git (`git clone https://github.com/esotericsoftware/spine-runtimes`) or download it as a zip via the download button above.
 3. If you are using C, add the sources from `spine-c/spine-c/src/spine` and `spine-sdl/src/spine-sdl-c.c` to your project, and add the folder `spine-c/spine-c/include` and `spine-sdl/src/spine-sdl-c.h` to your header search path. Note that includes are specified as `#inclue <spine/file.h>`, so the `spine` directory cannot be omitted when copying the source files.
-3. If you are using C++, add the sources from `spine-cpp/spine-cpp/src/spine` and `spine-sdl/src/spine-sdl-cpp.cpp` to your project, and add the folder `spine-cpp/spine-cpp/include` and `spine-sdl/src/spine-sdl-cpp.h` to your header search path. Note that includes are specified as `#inclue <spine/file.h>`, so the `spine` directory cannot be omitted when copying the source files.
+3. If you are using C++, add the sources from `spine-cpp/spine-cpp/src/spine` and `spine-sdl/src/spine-sdl-cpp.cpp` to your project, and add the folder `spine-cpp/spine-cpp/include` and `spine-sdl/src/spine-sdl-cpp.h` to your header search path. Note that includes are specified as `#include <spine/file.h>`, so the `spine` directory cannot be omitted when copying the source files.
 
 See the [Spine Runtimes documentation](http://esotericsoftware.com/spine-documentation#runtimesTitle) on how to use the APIs or check out the Spine SDL example in this repository.
 
diff --git a/spine-sdl/src/spine-sdl-cpp.cpp b/spine-sdl/src/spine-sdl-cpp.cpp
index c7c7c90a6..9cbba5504 100644
--- a/spine-sdl/src/spine-sdl-cpp.cpp
+++ b/spine-sdl/src/spine-sdl-cpp.cpp
@@ -114,8 +114,8 @@ void SkeletonDrawable::draw(SDL_Renderer *renderer) {
 			}
 
 			worldVertices.setSize(mesh->getWorldVerticesLength(), 0);
-			texture = (SDL_Texture *) mesh->getRegion()->rendererObject;
 			mesh->computeWorldVertices(slot, 0, mesh->getWorldVerticesLength(), worldVertices.buffer(), 0, 2);
+			texture = (SDL_Texture *) ((AtlasRegion *) mesh->getRendererObject())->page->getRendererObject();
 			verticesCount = mesh->getWorldVerticesLength() >> 1;
 			uvs = &mesh->getUVs();
 			indices = &mesh->getTriangles();
@@ -221,4 +221,4 @@ void SDLTextureLoader::unload(void *texture) {
 
 SpineExtension *spine::getDefaultExtension() {
 	return new DefaultSpineExtension();
-}
\ No newline at end of file
+}
diff --git a/spine-sfml/c/src/spine/spine-sfml.cpp b/spine-sfml/c/src/spine/spine-sfml.cpp
index 71615b672..56d79fe14 100644
--- a/spine-sfml/c/src/spine/spine-sfml.cpp
+++ b/spine-sfml/c/src/spine/spine-sfml.cpp
@@ -211,12 +211,12 @@ namespace spine {
 				}
 
 				if (mesh->super.worldVerticesLength > SPINE_MESH_VERTEX_COUNT_MAX) continue;
-				texture = (Texture *) ((spAtlasRegion *) mesh->rendererObject)->page->rendererObject;
 				spVertexAttachment_computeWorldVertices(SUPER(mesh), slot, 0, mesh->super.worldVerticesLength, worldVertices, 0, 2);
 				verticesCount = mesh->super.worldVerticesLength >> 1;
 				uvs = mesh->uvs;
 				indices = mesh->triangles;
 				indicesCount = mesh->trianglesCount;
+				texture = (Texture *) ((spAtlasRegion *) mesh->rendererObject)->page->rendererObject;
 
 			} else if (attachment->type == SP_ATTACHMENT_CLIPPING) {
 				spClippingAttachment *clip = (spClippingAttachment *) slot->attachment;
diff --git a/spine-sfml/cpp/src/spine/spine-sfml.cpp b/spine-sfml/cpp/src/spine/spine-sfml.cpp
index 50c08fe60..8b98c2a03 100644
--- a/spine-sfml/cpp/src/spine/spine-sfml.cpp
+++ b/spine-sfml/cpp/src/spine/spine-sfml.cpp
@@ -136,11 +136,11 @@ namespace spine {
 				}
 
 				worldVertices.setSize(mesh->getWorldVerticesLength(), 0);
-				texture = (Texture *) mesh->getRegion()->rendererObject;
 				mesh->computeWorldVertices(slot, 0, mesh->getWorldVerticesLength(), worldVertices.buffer(), 0, 2);
 				uvs = &mesh->getUVs();
 				indices = &mesh->getTriangles();
 				indicesCount = mesh->getTriangles().size();
+				texture = (Texture *) ((AtlasRegion *) mesh->getRendererObject())->page->getRendererObject();
 
 			} else if (attachment->getRTTI().isExactly(ClippingAttachment::rtti)) {
 				ClippingAttachment *clip = (ClippingAttachment *) slot.getAttachment();
diff --git a/spine-ts/spine-canvas/src/SkeletonRenderer.ts b/spine-ts/spine-canvas/src/SkeletonRenderer.ts
index 67340c5bd..55b73bbbd 100644
--- a/spine-ts/spine-canvas/src/SkeletonRenderer.ts
+++ b/spine-ts/spine-canvas/src/SkeletonRenderer.ts
@@ -30,6 +30,8 @@
 import { Utils, Color, Skeleton, RegionAttachment, TextureAtlasRegion, BlendMode, MeshAttachment, Slot } from "@esotericsoftware/spine-core";
 import { CanvasTexture } from "./CanvasTexture";
 
+const worldVertices = Utils.newFloatArray(8);
+
 export class SkeletonRenderer {
 	static QUAD_TRIANGLES = [0, 1, 2, 2, 3, 0];
 	static VERTEX_SIZE = 2 + 2 + 4;
@@ -65,6 +67,7 @@ export class SkeletonRenderer {
 
 			let attachment = slot.getAttachment();
 			if (!(attachment instanceof RegionAttachment)) continue;
+			attachment.computeWorldVertices(slot, worldVertices, 0, 2);
 			let region: TextureAtlasRegion = <TextureAtlasRegion>attachment.region;
 			let image: HTMLImageElement = (<CanvasTexture>region.page.texture).getImage() as HTMLImageElement;
 
diff --git a/spine-ts/spine-core/src/attachments/RegionAttachment.ts b/spine-ts/spine-core/src/attachments/RegionAttachment.ts
index 598bcfda9..6c46306a2 100644
--- a/spine-ts/spine-core/src/attachments/RegionAttachment.ts
+++ b/spine-ts/spine-core/src/attachments/RegionAttachment.ts
@@ -88,6 +88,20 @@ export class RegionAttachment extends Attachment implements HasTextureRegion {
 	updateRegion (): void {
 		if (!this.region) throw new Error("Region not set.");
 		let region = this.region;
+		let uvs = this.uvs;
+
+		if (region == null) {
+			uvs[0] = 0;
+			uvs[1] = 0;
+			uvs[2] = 0;
+			uvs[3] = 1;
+			uvs[4] = 1;
+			uvs[5] = 1;
+			uvs[6] = 1;
+			uvs[7] = 0;
+			return;
+		}
+
 		let regionScaleX = this.width / this.region.originalWidth * this.scaleX;
 		let regionScaleY = this.height / this.region.originalHeight * this.scaleY;
 		let localX = -this.width / 2 * this.scaleX + this.region.offsetX * regionScaleX;
@@ -116,16 +130,15 @@ export class RegionAttachment extends Attachment implements HasTextureRegion {
 		offset[6] = localX2Cos - localYSin;
 		offset[7] = localYCos + localX2Sin;
 
-		let uvs = this.uvs;
 		if (region.degrees == 90) {
+			uvs[0] = region.u2;
+			uvs[1] = region.v2;
 			uvs[2] = region.u;
 			uvs[3] = region.v2;
 			uvs[4] = region.u;
 			uvs[5] = region.v;
 			uvs[6] = region.u2;
 			uvs[7] = region.v;
-			uvs[0] = region.u2;
-			uvs[1] = region.v2;
 		} else {
 			uvs[0] = region.u;
 			uvs[1] = region.v2;
diff --git a/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SSpineWidget.cpp b/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SSpineWidget.cpp
index 4a845f1f1..90b9d29e2 100644
--- a/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SSpineWidget.cpp
+++ b/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SSpineWidget.cpp
@@ -274,8 +274,8 @@ void SSpineWidget::UpdateMesh(int32 LayerId, FSlateWindowElementList &OutDrawEle
 		if (attachment->getRTTI().isExactly(RegionAttachment::rtti)) {
 			RegionAttachment *regionAttachment = (RegionAttachment *) attachment;
 			attachmentColor.set(regionAttachment->getColor());
-			attachmentAtlasRegion = (AtlasRegion *) regionAttachment->getRendererObject();
 			regionAttachment->computeWorldVertices(*slot, *attachmentVertices, 0, 2);
+			attachmentAtlasRegion = (AtlasRegion *) regionAttachment->getRendererObject();
 			attachmentIndices = quadIndices;
 			attachmentUvs = regionAttachment->getUVs().buffer();
 			numVertices = 4;
@@ -283,8 +283,8 @@ void SSpineWidget::UpdateMesh(int32 LayerId, FSlateWindowElementList &OutDrawEle
 		} else if (attachment->getRTTI().isExactly(MeshAttachment::rtti)) {
 			MeshAttachment *mesh = (MeshAttachment *) attachment;
 			attachmentColor.set(mesh->getColor());
-			attachmentAtlasRegion = (AtlasRegion *) mesh->getRendererObject();
 			mesh->computeWorldVertices(*slot, 0, mesh->getWorldVerticesLength(), attachmentVertices->buffer(), 0, 2);
+			attachmentAtlasRegion = (AtlasRegion *) mesh->getRendererObject();
 			attachmentIndices = mesh->getTriangles().buffer();
 			attachmentUvs = mesh->getUVs().buffer();
 			numVertices = mesh->getWorldVerticesLength() >> 1;
diff --git a/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SpineSkeletonRendererComponent.cpp b/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SpineSkeletonRendererComponent.cpp
index 7b9eeedba..022079d14 100644
--- a/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SpineSkeletonRendererComponent.cpp
+++ b/spine-ue4/Plugins/SpinePlugin/Source/SpinePlugin/Private/SpineSkeletonRendererComponent.cpp
@@ -236,8 +236,8 @@ void USpineSkeletonRendererComponent::UpdateMesh(Skeleton *Skeleton) {
 			}
 
 			attachmentColor.set(regionAttachment->getColor());
-			attachmentAtlasRegion = (AtlasRegion *) regionAttachment->getRendererObject();
 			regionAttachment->computeWorldVertices(*slot, *attachmentVertices, 0, 2);
+			attachmentAtlasRegion = (AtlasRegion *) regionAttachment->getRendererObject();
 			attachmentIndices = quadIndices;
 			attachmentUvs = regionAttachment->getUVs().buffer();
 			numVertices = 4;
@@ -252,8 +252,8 @@ void USpineSkeletonRendererComponent::UpdateMesh(Skeleton *Skeleton) {
 			}
 
 			attachmentColor.set(mesh->getColor());
-			attachmentAtlasRegion = (AtlasRegion *) mesh->getRendererObject();
 			mesh->computeWorldVertices(*slot, 0, mesh->getWorldVerticesLength(), attachmentVertices->buffer(), 0, 2);
+			attachmentAtlasRegion = (AtlasRegion *) mesh->getRendererObject();
 			attachmentIndices = mesh->getTriangles().buffer();
 			attachmentUvs = mesh->getUVs().buffer();
 			numVertices = mesh->getWorldVerticesLength() >> 1;