diff --git a/spine-cocos2dx/src/spine/SkeletonRenderer.cpp b/spine-cocos2dx/src/spine/SkeletonRenderer.cpp
index e3035e23e..bc6c8cca4 100644
--- a/spine-cocos2dx/src/spine/SkeletonRenderer.cpp
+++ b/spine-cocos2dx/src/spine/SkeletonRenderer.cpp
@@ -282,8 +282,10 @@ void SkeletonRenderer::draw (Renderer* renderer, const Mat4& transform, uint32_t
 		
 		color.a *= nodeColor.a * _skeleton->color.a * slot->color.a * 255;
 		// skip rendering if the color of this attachment is 0
-		if (color.a == 0)
+		if (color.a == 0){
+			spSkeletonClipping_clipEnd(_clipper, slot);
 			continue;
+		}
 		float multiplier = _premultipliedAlpha ? color.a : 255;
 		color.r *= nodeColor.r * _skeleton->color.r * slot->color.r * multiplier;
 		color.g *= nodeColor.g * _skeleton->color.g * slot->color.g * multiplier;
@@ -313,8 +315,10 @@ void SkeletonRenderer::draw (Renderer* renderer, const Mat4& transform, uint32_t
 				spSkeletonClipping_clipTriangles(_clipper, (float*)&triangles.verts[0].vertices, triangles.vertCount * sizeof(cocos2d::V3F_C4B_T2F) / 4, triangles.indices, triangles.indexCount, (float*)&triangles.verts[0].texCoords, 6);
 				batch->deallocateVertices(triangles.vertCount);
 				
-				if (_clipper->clippedTriangles->size == 0)
+				if (_clipper->clippedTriangles->size == 0){
+					spSkeletonClipping_clipEnd(_clipper, slot);
 					continue;
+				}
 				
 				triangles.vertCount = _clipper->clippedVertices->size >> 1;
 				triangles.verts = batch->allocateVertices(triangles.vertCount);
@@ -397,8 +401,10 @@ void SkeletonRenderer::draw (Renderer* renderer, const Mat4& transform, uint32_t
 				spSkeletonClipping_clipTriangles(_clipper, (float*)&trianglesTwoColor.verts[0].position, trianglesTwoColor.vertCount * sizeof(V3F_C4B_C4B_T2F) / 4, trianglesTwoColor.indices, trianglesTwoColor.indexCount, (float*)&trianglesTwoColor.verts[0].texCoords, 7);
 				twoColorBatch->deallocateVertices(trianglesTwoColor.vertCount);
 				
-				if (_clipper->clippedTriangles->size == 0)
+				if (_clipper->clippedTriangles->size == 0){
+					spSkeletonClipping_clipEnd(_clipper, slot);
 					continue;
+				}
 				
 				trianglesTwoColor.vertCount = _clipper->clippedVertices->size >> 1;
 				trianglesTwoColor.verts = twoColorBatch->allocateVertices(trianglesTwoColor.vertCount);
diff --git a/spine-csharp/src/AnimationState.cs b/spine-csharp/src/AnimationState.cs
index 770fc91e7..2f7a47bc2 100644
--- a/spine-csharp/src/AnimationState.cs
+++ b/spine-csharp/src/AnimationState.cs
@@ -34,21 +34,21 @@ using System.Collections.Generic;
 namespace Spine {
 	public class AnimationState {
 		static readonly Animation EmptyAnimation = new Animation("<empty>", new ExposedList<Timeline>(), 0);
-		internal const int SUBSEQUENT = 0, FIRST = 1, DIP = 2, DIP_MIX = 3;
+		internal const int Subsequent = 0, First = 1, Dip = 2, DipMix = 3;
 
 		private AnimationStateData data;
-		private readonly ExposedList<TrackEntry> tracks = new ExposedList<TrackEntry>();
-		private readonly HashSet<int> propertyIDs = new HashSet<int>();
-		private readonly ExposedList<Event> events = new ExposedList<Event>();
-		private readonly EventQueue queue;
 
+		Pool<TrackEntry> trackEntryPool = new Pool<TrackEntry>();
+		private readonly ExposedList<TrackEntry> tracks = new ExposedList<TrackEntry>();
+		private readonly ExposedList<Event> events = new ExposedList<Event>();
+		private readonly EventQueue queue; // Initialized by constructor.
+
+		private readonly HashSet<int> propertyIDs = new HashSet<int>();
 		private readonly ExposedList<TrackEntry> mixingTo = new ExposedList<TrackEntry>();
 		private bool animationsChanged;
 
 		private float timeScale = 1;
 
-		Pool<TrackEntry> trackEntryPool = new Pool<TrackEntry>();
-
 		public AnimationStateData Data { get { return data; } }
 		/// <summary>A list of tracks that have animations, which may contain nulls.</summary>
 		public ExposedList<TrackEntry> Tracks { get { return tracks; } }
@@ -63,13 +63,13 @@ namespace Spine {
 		public AnimationState (AnimationStateData data) {
 			if (data == null) throw new ArgumentNullException("data", "data cannot be null.");
 			this.data = data;
-			this.queue = new EventQueue(this, HandleAnimationsChanged, trackEntryPool);
+			this.queue = new EventQueue(
+				this,
+				delegate { this.animationsChanged = true; },
+				trackEntryPool
+			);
 		}
-
-		void HandleAnimationsChanged () {
-			this.animationsChanged = true;
-		}
-
+			
 		/// <summary>
 		/// Increments the track entry times, setting queued animations as current if needed</summary>
 		/// <param name="delta">delta time</param>
@@ -155,7 +155,6 @@ namespace Spine {
 			return false;
 		}
 
-
 		/// <summary>
 		/// Poses the skeleton using the track entry animations. There are no side effects other than invoking listeners, so the 
 		/// animation state can be applied to multiple skeletons to pose them identically.</summary>
@@ -197,7 +196,7 @@ namespace Spine {
 
 					for (int ii = 0; ii < timelineCount; ii++) {
 						Timeline timeline = timelinesItems[ii];
-						MixPose pose = timelineData[ii] >= FIRST ? MixPose.Setup : currentPose;
+						MixPose pose = timelineData[ii] >= AnimationState.First ? MixPose.Setup : currentPose;
 						var rotateTimeline = timeline as RotateTimeline;
 						if (rotateTimeline != null)
 							ApplyRotateTimeline(rotateTimeline, skeleton, animationTime, mix, pose, timelinesRotation, ii << 1, firstFrame);
@@ -246,17 +245,17 @@ namespace Spine {
 			for (int i = 0; i < timelineCount; i++) {
 				Timeline timeline = timelinesItems[i];
 				switch (timelineData[i]) {
-				case SUBSEQUENT:
+				case Subsequent:
 					if (!attachments && timeline is AttachmentTimeline) continue;
 					if (!drawOrder && timeline is DrawOrderTimeline) continue;
 					pose = currentPose;
 					alpha = alphaMix;
 					break;
-				case FIRST:
+				case First:
 					pose = MixPose.Setup;
 					alpha = alphaMix;
 					break;
-				case DIP:
+				case Dip:
 					pose = MixPose.Setup;
 					alpha = alphaDip;
 					break;
@@ -421,6 +420,7 @@ namespace Spine {
 			queue.Drain();
 		}
 
+		/// <summary>Sets the active TrackEntry for a given track number.</summary>
 		private void SetCurrent (int index, TrackEntry current, bool interrupt) {
 			TrackEntry from = ExpandToIndex(index);
 			tracks.Items[index] = current;
@@ -437,12 +437,12 @@ namespace Spine {
 				from.timelinesRotation.Clear(); // Reset rotation for mixing out, in case entry was mixed in.
 			}
 
-			queue.Start(current);
+			queue.Start(current); // triggers AnimationsChanged
 		}
 
 
 		/// <summary>Sets an animation by name. <seealso cref="SetAnimation(int, Animation, bool)" /></summary>
-		public TrackEntry SetAnimation (int trackIndex, String animationName, bool loop) {
+		public TrackEntry SetAnimation (int trackIndex, string animationName, bool loop) {
 			Animation animation = data.skeletonData.FindAnimation(animationName);
 			if (animation == null) throw new ArgumentException("Animation not found: " + animationName, "animationName");
 			return SetAnimation(trackIndex, animation, loop);
@@ -480,7 +480,7 @@ namespace Spine {
 
 		/// <summary>Queues an animation by name.</summary>
 		/// <seealso cref="AddAnimation(int, Animation, bool, float)" />
-		public TrackEntry AddAnimation (int trackIndex, String animationName, bool loop, float delay) {
+		public TrackEntry AddAnimation (int trackIndex, string animationName, bool loop, float delay) {
 			Animation animation = data.skeletonData.FindAnimation(animationName);
 			if (animation == null) throw new ArgumentException("Animation not found: " + animationName, "animationName");
 			return AddAnimation(trackIndex, animation, loop, delay);
@@ -570,6 +570,7 @@ namespace Spine {
 			return null;
 		}
 
+		/// <summary>Object-pooling version of new TrackEntry. Obtain an unused TrackEntry from the pool and clear/initialize its values.</summary>
 		/// <param name="last">May be null.</param>
 		private TrackEntry NewTrackEntry (int trackIndex, Animation animation, bool loop, TrackEntry last) {
 			TrackEntry entry = trackEntryPool.Obtain(); // Pooling
@@ -589,7 +590,7 @@ namespace Spine {
 			entry.delay = 0;
 			entry.trackTime = 0;
 			entry.trackLast = -1;
-			entry.nextTrackLast = -1;
+			entry.nextTrackLast = -1; // nextTrackLast == -1 signifies a TrackEntry that wasn't applied yet.
 			entry.trackEnd = float.MaxValue; // loop ? float.MaxValue : animation.Duration;
 			entry.timeScale = 1;
 
@@ -600,6 +601,7 @@ namespace Spine {
 			return entry;
 		}
 
+		/// <summary>Dispose all track entries queued after the given TrackEntry.</summary>
 		private void DisposeNext (TrackEntry entry) {
 			TrackEntry next = entry.next;
 			while (next != null) {
@@ -628,7 +630,7 @@ namespace Spine {
 			return (trackIndex >= tracks.Count) ? null : tracks.Items[trackIndex];
 		}
 
-		override public String ToString () {
+		override public string ToString () {
 			var buffer = new System.Text.StringBuilder();
 			for (int i = 0, n = tracks.Count; i < n; i++) {
 				TrackEntry entry = tracks.Items[i];
@@ -680,11 +682,12 @@ namespace Spine {
 			Event = null;
 		}
 
+		/// <summary>Sets the timeline data.</summary>
 		/// <param name="to">May be null.</param>
 		internal TrackEntry SetTimelineData (TrackEntry to, ExposedList<TrackEntry> mixingToArray, HashSet<int> propertyIDs) {
 			if (to != null) mixingToArray.Add(to);
 			var lastEntry = mixingFrom != null ? mixingFrom.SetTimelineData(this, mixingToArray, propertyIDs) : this;
-			if (to != null) mixingToArray.RemoveAt(mixingToArray.Count - 1); // mixingToArray.pop();
+			if (to != null) mixingToArray.Pop();
 
 			var mixingTo = mixingToArray.Items;
 			int mixingToLast = mixingToArray.Count - 1;
@@ -698,21 +701,19 @@ namespace Spine {
 			for (int i = 0; i < timelinesCount; i++) {
 				int id = timelines[i].PropertyId;
 				if (!propertyIDs.Add(id)) {
-					timelineDataItems[i] = AnimationState.SUBSEQUENT;
+					timelineDataItems[i] = AnimationState.Subsequent;
 				} else if (to == null || !to.HasTimeline(id)) {
-					timelineDataItems[i] = AnimationState.FIRST;
+					timelineDataItems[i] = AnimationState.First;
 				} else {
 					for (int ii = mixingToLast; ii >= 0; ii--) {
 						var entry = mixingTo[ii];
-						if (!entry.HasTimeline(id)) {
-							if (entry.mixDuration > 0) {
-								timelineDataItems[i] = AnimationState.DIP_MIX;
-								timelineDipMixItems[i] = entry;
-								goto outer; // continue outer;
-							}
+						if (entry.mixDuration > 0 && !entry.HasTimeline(id)) {
+							timelineDataItems[i] = AnimationState.DipMix;
+							timelineDipMixItems[i] = entry;
+							goto outer; // continue outer;
 						}
 					}
-					timelineDataItems[i] = AnimationState.DIP;
+					timelineDataItems[i] = AnimationState.Dip;
 				}
 				outer: {}
 			}
@@ -886,20 +887,20 @@ namespace Spine {
 			timelinesRotation.Clear();
 		}
 
-		override public String ToString () {
+		override public string ToString () {
 			return animation == null ? "<none>" : animation.name;
 		}
 	}
 
 	class EventQueue {
 		private readonly List<EventQueueEntry> eventQueueEntries = new List<EventQueueEntry>();
-		public bool drainDisabled;
+		internal bool drainDisabled;
 
 		private readonly AnimationState state;
 		private readonly Pool<TrackEntry> trackEntryPool;
-		public event Action AnimationsChanged;
+		internal event Action AnimationsChanged;
 
-		public EventQueue (AnimationState state, Action HandleAnimationsChanged, Pool<TrackEntry> trackEntryPool) {
+		internal EventQueue (AnimationState state, Action HandleAnimationsChanged, Pool<TrackEntry> trackEntryPool) {
 			this.state = state;
 			this.AnimationsChanged += HandleAnimationsChanged;
 			this.trackEntryPool = trackEntryPool;
@@ -921,33 +922,34 @@ namespace Spine {
 			Start, Interrupt, End, Dispose, Complete, Event
 		}
 
-		public void Start (TrackEntry entry) {
+		internal void Start (TrackEntry entry) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.Start, entry));
 			if (AnimationsChanged != null) AnimationsChanged();
 		}
 
-		public void Interrupt (TrackEntry entry) {
+		internal void Interrupt (TrackEntry entry) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.Interrupt, entry));
 		}
 
-		public void End (TrackEntry entry) {
+		internal void End (TrackEntry entry) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.End, entry));
 			if (AnimationsChanged != null) AnimationsChanged();
 		}
 
-		public void Dispose (TrackEntry entry) {
+		internal void Dispose (TrackEntry entry) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.Dispose, entry));
 		}
 
-		public void Complete (TrackEntry entry) {
+		internal void Complete (TrackEntry entry) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.Complete, entry));
 		}
 
-		public void Event (TrackEntry entry, Event e) {
+		internal void Event (TrackEntry entry, Event e) {
 			eventQueueEntries.Add(new EventQueueEntry(EventType.Event, entry, e));
 		}
 
-		public void Drain () {
+		/// <summary>Raises all events in the queue and drains the queue.</summary>
+		internal void Drain () {
 			if (drainDisabled) return;
 			drainDisabled = true;
 
@@ -992,7 +994,7 @@ namespace Spine {
 			drainDisabled = false;
 		}
 
-		public void Clear () {
+		internal void Clear () {
 			eventQueueEntries.Clear();
 		}
 	}
diff --git a/spine-csharp/src/Atlas.cs b/spine-csharp/src/Atlas.cs
index 04f2c5755..b58897153 100644
--- a/spine-csharp/src/Atlas.cs
+++ b/spine-csharp/src/Atlas.cs
@@ -43,11 +43,21 @@ using Windows.Storage;
 #endif
 
 namespace Spine {
-	public class Atlas {
+	public class Atlas : IEnumerable<AtlasRegion> {
 		readonly List<AtlasPage> pages = new List<AtlasPage>();
 		List<AtlasRegion> regions = new List<AtlasRegion>();
 		TextureLoader textureLoader;
 
+		#region IEnumerable implementation
+		public IEnumerator<AtlasRegion> GetEnumerator () {
+			return regions.GetEnumerator();
+		}
+
+		System.Collections.IEnumerator System.Collections.IEnumerable.GetEnumerator () {
+			return regions.GetEnumerator();
+		}
+		#endregion
+
 		#if !(IS_UNITY)
 		#if WINDOWS_STOREAPP
 		private async Task ReadFile(string path, TextureLoader textureLoader) {
@@ -99,7 +109,7 @@ namespace Spine {
 		}
 
 		private void Load (TextReader reader, string imagesDir, TextureLoader textureLoader) {
-			if (textureLoader == null) throw new ArgumentNullException("textureLoader cannot be null.");
+			if (textureLoader == null) throw new ArgumentNullException("textureLoader", "textureLoader cannot be null.");
 			this.textureLoader = textureLoader;
 
 			string[] tuple = new string[4];
@@ -168,11 +178,11 @@ namespace Spine {
 					region.height = Math.Abs(height);
 
 					if (ReadTuple(reader, tuple) == 4) { // split is optional
-						region.splits = new int[] {int.Parse(tuple[0]), int.Parse(tuple[1]),
+						region.splits = new [] {int.Parse(tuple[0]), int.Parse(tuple[1]),
 								int.Parse(tuple[2]), int.Parse(tuple[3])};
 
 						if (ReadTuple(reader, tuple) == 4) { // pad is optional, but only present with splits
-							region.pads = new int[] {int.Parse(tuple[0]), int.Parse(tuple[1]),
+							region.pads = new [] {int.Parse(tuple[0]), int.Parse(tuple[1]),
 									int.Parse(tuple[2]), int.Parse(tuple[3])};
 
 							ReadTuple(reader, tuple);
diff --git a/spine-csharp/src/ExposedList.cs b/spine-csharp/src/ExposedList.cs
index 421c6a2c3..c2c7de4d9 100644
--- a/spine-csharp/src/ExposedList.cs
+++ b/spine-csharp/src/ExposedList.cs
@@ -104,14 +104,14 @@ namespace Spine {
 			}
 		}
 
-		private void CheckRange (int idx, int count) {
-			if (idx < 0)
+		private void CheckRange (int index, int count) {
+			if (index < 0)
 				throw new ArgumentOutOfRangeException("index");
 
 			if (count < 0)
 				throw new ArgumentOutOfRangeException("count");
 
-			if ((uint)idx + (uint)count > (uint)Count)
+			if ((uint)index + (uint)count > (uint)Count)
 				throw new ArgumentException("index and count exceed length of list");
 		}
 
@@ -450,6 +450,21 @@ namespace Spine {
 			version++;
 		}
 
+		// Spine Added Method
+		// Based on Stack<T>.Pop(); https://referencesource.microsoft.com/#mscorlib/system/collections/stack.cs
+		/// <summary>Pops the last item of the list. If the list is empty, Pop throws an InvalidOperationException.</summary>
+		public T Pop () {
+			if (Count == 0)
+				throw new InvalidOperationException("List is empty. Nothing to pop.");
+			
+			int i = Count - 1;
+			T item = Items[i];
+			Items[i] = default(T);
+			Count--;
+			version++;
+			return item;
+		}
+
 		public void RemoveRange (int index, int count) {
 			CheckRange(index, count);
 			if (count > 0) {
diff --git a/spine-starling/spine-starling-example/lib/spine-starling.swc b/spine-starling/spine-starling-example/lib/spine-starling.swc
index 0774a336f..91ad4b906 100644
Binary files a/spine-starling/spine-starling-example/lib/spine-starling.swc and b/spine-starling/spine-starling-example/lib/spine-starling.swc differ
diff --git a/spine-starling/spine-starling/src/spine/starling/StarlingAtlasAttachmentLoader.as b/spine-starling/spine-starling/src/spine/starling/StarlingAtlasAttachmentLoader.as
index 7c6543381..62467744e 100644
--- a/spine-starling/spine-starling/src/spine/starling/StarlingAtlasAttachmentLoader.as
+++ b/spine-starling/spine-starling/src/spine/starling/StarlingAtlasAttachmentLoader.as
@@ -50,7 +50,6 @@ package spine.starling {
 
 		public function StarlingAtlasAttachmentLoader(atlas : TextureAtlas) {
 			this.atlas = atlas;
-
 			Bone.yDown = true;
 		}
 
@@ -59,11 +58,11 @@ package spine.starling {
 		}
 
 		public function newRegionAttachment(skin : Skin, name : String, path : String) : RegionAttachment {
-			var texture : Texture = getTexture(path);
+			var texture : SubTexture = getTexture(path) as SubTexture;
 			if (texture == null)
 				throw new Error("Region not found in Starling atlas: " + path + " (region attachment: " + name + ")");
 			var attachment : RegionAttachment = new RegionAttachment(name);
-			var rotated : Boolean = atlas.getRotation(path);
+			var rotated : Boolean = texture.rotated;
 			attachment.rendererObject = new Image(Texture.fromTexture(texture)); // Discard frame.
 			var frame : Rectangle = texture.frame;
 			attachment.regionOffsetX = frame ? -frame.x : 0;
@@ -79,59 +78,44 @@ package spine.starling {
 				tmp = attachment.regionWidth;
 				attachment.regionWidth = attachment.regionHeight;
 				attachment.regionHeight = tmp;
+                attachment["regionU2"] = 0;
+                attachment["regionV2"] = 1;
+                attachment["regionU"] = 1;
+                attachment["regionV"] = 0;
+			}else{
+                attachment["regionU"] = 0;
+                attachment["regionV"] = 0;
+                attachment["regionU2"] = 1;
+                attachment["regionV2"] = 1;
 			}
-			if (!rotated) {
-				attachment["regionU"] = 0;
-				attachment["regionV"] = 0;
-				attachment["regionU2"] = 1;
-				attachment["regionV2"] = 1;
-			} else {
-				attachment["regionU2"] = 0;
-				attachment["regionV2"] = 1;
-				attachment["regionU"] = 1;
-				attachment["regionV"] = 0;
-			}
-			attachment.setUVs(attachment["regionU"], attachment["regionV"], attachment["regionU2"], attachment["regionV2"], atlas.getRotation(path));
+			attachment.setUVs(attachment["regionU"], attachment["regionV"], attachment["regionU2"], attachment["regionV2"], rotated);
 			return attachment;
 		}
 
 		public function newMeshAttachment(skin : Skin, name : String, path : String) : MeshAttachment {
-			var texture : Texture = getTexture(path);
+			var texture : SubTexture = getTexture(path) as SubTexture;
 			if (texture == null)
 				throw new Error("Region not found in Starling atlas: " + path + " (mesh attachment: " + name + ")");
-			var rotated : Boolean = atlas.getRotation(path);
+			var rotated : Boolean = texture.rotated;
 			var attachment : MeshAttachment = new MeshAttachment(name);
 			attachment.regionRotate = rotated;
 			attachment.rendererObject = new Image(Texture.fromTexture(texture)); // Discard frame.
-			var subTexture : SubTexture = texture as SubTexture;
-			if (subTexture) {
-				var root : Texture = subTexture.root;
-				var rectRegion : Rectangle = atlas.getRegion(path);
-				if (!rotated) {
-					attachment.regionU = rectRegion.x / root.width;
-					attachment.regionV = rectRegion.y / root.height;
-					attachment.regionU2 = (rectRegion.x + subTexture.width) / root.width;
-					attachment.regionV2 = (rectRegion.y + subTexture.height) / root.height;
-				} else {
-					attachment.regionU2 = rectRegion.x / root.width;
-					attachment.regionV2 = rectRegion.y / root.height;
-					attachment.regionU = (rectRegion.x + subTexture.height) / root.width;
-					attachment.regionV = (rectRegion.y + subTexture.width) / root.height;
-				}
-				attachment.rendererObject = new Image(root);
+
+			var root : Texture = texture.root;
+			var rectRegion : Rectangle = atlas.getRegion(path);
+			if (!rotated) {
+				attachment.regionU = rectRegion.x / root.width;
+				attachment.regionV = rectRegion.y / root.height;
+				attachment.regionU2 = (rectRegion.x + texture.width) / root.width;
+				attachment.regionV2 = (rectRegion.y + texture.height) / root.height;
 			} else {
-				if (!rotated) {
-					attachment.regionU = 0;
-					attachment.regionV = 1;
-					attachment.regionU2 = 1;
-					attachment.regionV2 = 0;
-				} else {
-					attachment.regionU2 = 0;
-					attachment.regionV2 = 1;
-					attachment.regionU = 1;
-					attachment.regionV = 0;
-				}
+				attachment.regionU2 = rectRegion.x / root.width;
+				attachment.regionV2 = rectRegion.y / root.height;
+				attachment.regionU = (rectRegion.x + texture.height) / root.width;
+				attachment.regionV = (rectRegion.y + texture.width) / root.height;
 			}
+			attachment.rendererObject = new Image(root);
+
 			var frame : Rectangle = texture.frame;
 			attachment.regionOffsetX = frame ? -frame.x : 0;
 			attachment.regionOffsetY = frame ? -frame.y : 0;
diff --git a/spine-unity/Assets/spine-unity/SkeletonAnimator.cs b/spine-unity/Assets/spine-unity/SkeletonAnimator.cs
index 7d76b28ac..0337eedc4 100644
--- a/spine-unity/Assets/spine-unity/SkeletonAnimator.cs
+++ b/spine-unity/Assets/spine-unity/SkeletonAnimator.cs
@@ -75,7 +75,12 @@ namespace Spine.Unity {
 		public void Update () {
 			if (!valid) return;
 
+			#if UNITY_EDITOR
+			if (Application.isPlaying)
+				translator.Apply(skeleton);
+			#else
 			translator.Apply(skeleton);
+			#endif
 
 			// UpdateWorldTransform and Bone Callbacks
 			{